1. Functional Morphology and Biomechanics, Institute of Zoology, Kiel University, Kiel 24118, Germany
2. Division of Mechanical Engineering and Design, School of Engineering, London South Bank University, London SE1 0AA, UK
3. Mechanical Intelligence Research Group, South Bank Applied BioEngineering Research (SABER) Centre, School of Engineering, London South Bank University, London SE1 0AA, UK