1. Department of Biological Sciences, Auburn University, Auburn, AL 36849, USA
2. Department of Biology and Integrated Bioscience Program, The University of Akron, Akron, OH 44325, USA
3. Department of Biology, Evolution and Optics of Nanostructures Group, University of Ghent, Ghent, Belgium
4. Department of Ecology, Evolution, and Organismal Biology, Iowa State University, Ames, IA 50011, USA