1. Group of Theoretical Biology, The State Key Laboratory of Bio-control, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, People’s Republic of China
2. Evolutionary Dynamics Group, Centre for Cancer Genomics and Computational Biology, Barts Cancer Institute, Queen Mary University of London, London, UK
3. The first affiliated hospital of Guangzhou Medical University, Guangzhou, People’s Republic of China
4. School of Mathematical Sciences, Queen Mary University of London, London, UK