1. Department of Life Sciences, Natural History Museum, London SW7 5BD, UK
2. Centre for Craniofacial and Regenerative Biology, King's College London SE1 9RT, UK
3. Division of Biosciences, University College London WC1E 6DE, UK
4. Univ Lyon, Université Claude Bernard Lyon 1, CNRS, ENTPE, UMR 5023 LEHNA, F-69622, Villeurbanne, France