1. Department of Mathematics, University of British Columbia1984 Mathematics Road, Vancouver, BC, Canada V6T 1Z2
2. Department of Earth and Ocean Sciences, University of British Columbia6339 Stores Road, Vancouver, BC, Canada V6T 1Z4
3. Department of Mathematics, Imperial College LondonSouth Kensington Campus, London SW7 2AZ, UK
4. Division of Mathematical Modeling, University of British Columbia Centre for Disease Control655 West 12th Avenue, Vancouver, BC, Canada V5Z 4R4