1. Department of Geography, Geology and Environment, University of Hull, Cottingham Road, Hull HU6 7RX, UK; Department of Earth Sciences, University College London, Gower Street, London WC1E 6BT, UK
2. Department of Palaeontology, Natural History Museum, Cromwell Road, London SW7 5BD, UK
3. Lansdowne, Military Road, Chale, Isle of Wight PO38 2HH, UK