1. Okinawa Institute of Science and Technology Graduate University, Onna-son, Okinawa 904-0495, Japan
2. National Institute of Genetics, Mishima, Shizuoka 411-8540, Japan
3. Graduate School of Biostudies, Kyoto University, Sakyo-ku, Kyoto 606-8501, Japan
4. Tokyo Institute of Technology, Graduate School of Bioscience and Biotechnology, Nagatsuda, Yokohama 226-8501, Japan