1. Chromosome Segregation Laboratory, The Francis Crick Institute, Lincoln's Inn Fields Laboratories, 44 Lincoln's Inn Fields, London WC2A 3LY, UK
2. Laboratory of Cytoskeletal Logistics, Department of Life Science and Medical Bioscience, Graduate School of Advanced Science and Engineering, Waseda University TWIns, 2-2 Wakamatsucho, Shinjuku, Tokyo 162-0056, Japan
3. Department of Biophysics and Biochemistry, Graduate School of Science, University of Tokyo, 7-3-1 Hongo, Tokyo 113-0033, Japan
4. The Gurdon Institute, University of Cambridge, Tennis Court Road, Cambridge CB2 1QN, UK
5. Department of Genetics, University of Cambridge, Downing Street, Cambridge CB2 3EH, UK
6. Department of Computational Biology, Research and Innovation Centre, Fondazione Edmund Mach, San Michele all’Adige 38010, Italy
7. Randall Division of Cell and Molecular Biophysics and Institute for Mathematical and Molecular Biomedicine, King's College London, London SE1 1UL, UK