1. Oxford Centre for Industrial and Applied Mathematics, Mathematical Institute, University of Oxford, Oxford OX2 6GG, UK
2. Computational Biology Laboratory, Fundación Ciencia & Vida, Avda. Zañartu 1482, Ñuñoa, Santiago, Chile
3. Department of Mathematics, University of California, Los Angeles, CA 90095, USA
4. Department of Mathematics and Statistics, University of Massachusetts, Amherst, MA 01003, USA