1. School of Earth and Environment, University of Leeds, Leeds LS2 9JT, UK
2. Geography, College of Life and Environmental Sciences, University of Exeter, Rennes Drive, Exeter EX4 4RJ, UK
3. School of Biology, University of Leeds, Leeds LS2 9JT, UK
4. Grantham Research Institute on Climate Change and the Environment, London School of Economics, London WC2A 2AE, UK
5. Climatic Research Unit, School of Environmental Sciences, University of East Anglia, Norwich Research Park, Norwich NR4 7TJ, UK
6. NZ Climate Change Research Institute, Victoria University, Wellington, PO Box 600, Wellington 6012, New Zealand