1. Green Templeton College, University of Oxford, Woodstock Road, Oxford, UK
2. Structural Biophysics Research Group, School of Optometry and Vision Sciences, Cardiff University, Maindy Road, Cardiff, UK
3. Clinical Investigation Group, School of Optometry and Vision Sciences, Cardiff University, Maindy Road, Cardiff, UK
4. Physical Sciences Group, DSTL Platform Systems Division, DSTL Porton Down, Porton Down, Salisbury, Wilts SP4 0JQ, UK