1. University of Bristol School of Biochemistry, Bristol BS8 1QU, UK
2. Genomics of Neurodegenerative Diseases and Aging, Vrije Universiteit Amsterdam, Amsterdam, The Netherlands
3. Department of Neurology, Columbia University, New York, NY 10032, USA
4. Department of Medicine (Division of Neurology), Temerty Faculty of Medicine, University Health Network, University of Toronto, Toronto, Canada, M5T 0S8