1. European Molecular Biology Laboratory, Heidelberg 69117, Germany
2. Computational Biology Group, Institute of Integrative Biology, Building CHN H 75.1, Universitätstrasse 16, CH-8092 Zurich, Switzerland
3. Swiss Institute of Bioinformatics, Quartier Sorge - Batiment Amphipole, 1015 Lausanne, Switzerland
4. Laboratoire de Physique de l'École Normale Supérieure, CNRS, PSL University, Sorbonne Université, and Université Paris Cité, 75005 Paris, France
5. Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109-1382, USA