1. Ecological Genetics Laboratory, National Institute of Genetics, Yata 1111, Mishima, Shizuoka 411-8540, Japan
2. Laboratory of Molecular Ecological Genetics, Graduate School of Frontier Sciences, The University of Tokyo, Kashiwanoha 5-1-5, Chiba 277-8562, Japan
3. School of Life Sciences, University of Nottingham, Nottingham NG7 2RD, UK
4. Institut Jacques Monod, CNRS UMR7592, Université Paris Cité, 15 rue Hélène Brion, 75013 Paris, France