1. ICREA-Complex Systems Lab, CSIC Universitat Pompeu Fabra, Dr Aiguader 80, Barcelona 08003, Spain
2. CSIC-UPF, Institut de Biologia Evolutiva, Pg Maritim de la Barceloneta 37, Barcelona 08003, Spain
3. Santa Fe Institute, 1399 Hyde Park Road, Santa Fe, NM 87501, USA
4. Department of Ecology and Evolutionary Biology, Princeton University, Princeton, NJ 08544, USA