1. College of Life Science, Shenyang Normal University, Shenyang, Liaoning 110034, People's Republic of China
2. Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, People's Republic of China
3. College of Resources and Environmental Sciences, Gansu Agricultural University, Lanzhou 730070, People's Republic of China
4. Life Science, Lanzhou University, Tianshui Road, Chengguan Qu, Lanzhou, Gansu Province 730000, People's Republic of China
5. Department of Anthropology, University College London, London WC1H 0BW, UK
6. IAST, Toulouse School of Economics, Toulouse, Occitanie, 31080, France