1. School of Natural Sciences, Bangor University, Bangor LL57 2DG, UK
2. School of BioSciences, University of Melbourne, Melbourne VIC 3010, Australia
3. Field Research Station at Fort Missoula, Division of Biological Sciences, University of Montana, MT 59802, USA
4. Lancaster Environment Centre, Lancaster University, Lancaster LA1 4YQ, UK
5. Department of Earth Sciences, Montana State University, Bozeman, MT 59717, USA
6. Department of Evolution, Ecology, and Behavior, School of Integrative Biology, University of Illinois at Urbana-Champaign, Urbana, IL 61801, USA