1. Department of Biology II, Ludwig-Maximilians University of Munich, Planegg-Martinsried, Munich, Germany
2. Institute of Zoology, University of Hamburg, Martin-Luther-King Platz 3, Hamburg 20146 Germany
3. General and Systematic Zoology, University of Greifswald, Bachstrasse 11/13, Greifswald 17489, Germany
4. Department of Biological Sciences, Macquarie University, North Ryde, Sydney NSW 2109, Australia