1. Department of Plant Protection Biology, Swedish University of Agricultural Sciences, PO Box 102, Alnarp 23053, Sweden
2. Royal Botanic Gardens, Kew, Kew Green, Richmond, Surrey TW9 3AE, UK
3. Natural Resources Institute, University of Greenwich, Chatham, Kent ME4 4TB, UK
4. Department of Botany, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland