1. Department of Marine Sciences, University of Gothenburg, 405 30 Gothenburg, Sweden
2. Centre for Marine Evolutionary Biology, University of Gothenburg, 405 30 Gothenburg, Sweden
3. Department of Environmental Systems Science, ETH Zürich, 8092 Zürich, Switzerland
4. Ecology and Evolutionary Biology, School of Biosciences, University of Sheffield, Sheffield S10 2TN, UK
5. Department of Marine Sciences, University of Gothenburg, Tjärnö Marine Laboratory, 452 96 Strömstad, Sweden