1. Institute of Biology, University of Neuchâtel, Neuchâtel, Switzerland
2. Budongo Conservation Field Station, Masindi, Uganda
3. School of Psychology and Neuroscience, University of St Andrews, St Andrews, UK
4. Centre for Research in Animal Behaviour, University of Exeter, Exeter, UK
5. Université Clermont Auvergne, LAPSCO CNRS, Clermont-Ferrand, France