1. Biosciences, Swansea University, Singleton Park, Swansea SA2 8PP, UK
2. Marine Mammal Research, Department of Bioscience, Aarhus University, Frederiksborgvej 399, 4000 Roskilde, Denmark
3. Institute for Terrestrial and Aquatic Wildlife Research, University of Veterinary Medicine Hannover, Bischofsholer Damm 15, 30173 Hannover, Germany