1. School of BioSciences, University of Melbourne, Parkville, Victoria 3010, Australia
2. Department of Ecology and Evolutionary Biology, Cornell University, Ithaca, NY 14853, USA
3. National Centre for Coasts and Climate (NCCC), School of BioSciences, University of Melbourne, Parkville, Victoria 3010, Australia
4. Centre for Marine Science and Innovation, Evolution and Ecology Research Centre, School of Biological, Earth and Environmental Science, University of New South Wales, Sydney, New South Wales 2052, Australia