1. Department of Biology, University of Oxford, Oxford OX1 3SZ, UK
2. School of Biological Sciences, Monash University, Melbourne 3800, Australia
3. Department of Evolutionary and Integrative Ecology, Leibniz Institute of Freshwater Ecology and Inland Fisheries (IGB), 12587 Berlin, Germany
4. Department of Biology, Chemistry, and Pharmacy, Institute of Biology, Freie Universität Berlin (FU), 14195 Berlin, Germany