1. Department of Biology, University of Oxford, Oxford OX1 3SZ, UK
2. Department of Zoology, Marine Apex Predator Research Unit, Institute for Coastal and Marine Research, Nelson Mandela University, Port Elizabeth-Gqeberha 6031, South Africa
3. Centre d'Etudes Biologiques de Chizé, UMR 7372 CNRS – La Rochelle Université, 79360 Villiers en Bois, France
4. Odum School of Ecology, University of Georgia, Athens, GA 30602, USA
5. Department of Brain and Cognitive Sciences, University of Rochester, Rochester, NY 14627, USA