1. International Research Center for Medical Sciences, Kumamoto University, Kumamoto 860-0811, Japan
2. Department of Physiology, Development and Neuroscience, University of Cambridge, Cambridge CB2 3EG, UK
3. Department of Organismal Biology and Anatomy, University of Chicago, Chicago, IL 60637, USA
4. Department of Physiology, Anatomy and Genetics, University of Oxford, Oxford OX3 7TY, UK