1. Eck Department of Biological Sciences and Eck Institute for Global Health, University of Notre Dame, Notre Dame, IN 46556, USA
2. Department of Epidemiology of Microbial Diseases, Yale School of Public Health, New Haven, CT 06510, USA
3. Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, CO 80523, USA