1. Department of Earth Sciences, University College London, Gower Street, London WC1E 6BT, UK
2. Department of Life Sciences, Natural History Museum, Cromwell Road, London SW7 5BD, UK
3. Science and Solutions for a Changing Planet DTP, and the Department of Life Sciences, Imperial College London, South Kensington Campus, London SW7 2AZ, UK
4. Laboratoire Géosystèmes, Environnement et Développement Durable, Département de Géologie, Faculté des Sciences Dhar El Mahraz, Université Sidi Mohamed Ben Abdellah, BP 1796, Atlas 30 000, Fès, Morocco