1. Centre for Complexity Science, Zeeman Building, University of Warwick, Coventry CV4 7AL, UK
2. School of Mathematics, University of Manchester, Oxford Road, Manchester M13 9PL, UK
3. Zeeman Institute: Systems Biology and Infectious Disease Epidemiology Research (SBIDER), University of Warwick, Coventry CV4 7AL, UK
4. Warwick Medical School, University of Warwick, Coventry CV4 7AL, UK
5. Centre for Health Policy, University of the Witwatersrand, Johannesburg, South Africa