1. Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, D-10115 Berlin, Germany
2. Department of Biological Sciences, University of of Alberta, Edmonton, Alberta T6G 2E9, Canada; Queensland Museum 4101 and University of the Sunshine Coast, Queensland, Australia, 4556
3. Plant Protection Institute, Centre for Agricultural Research, Hungarian Academy of Sciences, H-1525 Budapest, P.O. Box 102, Hungary