1. Department of Zoology & Entomology University of Pretoria, Private Bag X20 Hatfield, 0028, SOUTH AFRICA and Laboratório de Scarabaeoidologia, Instituto de Biociências Universidade Federal de Mato Grosso (UFMT) Cuiabá, Mato Grosso, BRAZIL https://orc
2. Canadian Museum of Nature 1740 Pink Road, Gatineau, QC J9J 3N7, CANADA https://orcid.org/0000-0002-8059-5133