1. 2Department of Ecology and Evolutionary Biology, University of California, Irvine, California 92697‐2525 USA;
2. 3School of Biological Sciences, University of Nebraska, Lincoln, Nebraska 68588‐0343 USA; and
3. 4Department of Ecology and Evolution, State University of New York at Stony Brook, Stony Brook, New York 11794‐5245 USA