1. Department of Biological Sciences, LSCB 124, University of South Alabama, Mobile, Alabama 36688 USA;
2. Department of Ecology and Evolutionary Biology, and Natural History Museum and
3. Biodiversity Research Center, University of Kansas, Lawrence, Kansas 66045 USA; and
4. Museo Paleontológico Egidio Feruglio, 9100 Trelew, Chubut, Argentina