1. National Herbarium Nederland, Wageningen University branch; and Biosystematics Group; Wageningen UR; Generaal Foulkesweg 37 6703 BL Wageningen The Netherlands
2. Department of Botany, Centre for Plant Diversity and Systematics; The University of Reading; P.O. Box 221, Reading RG6 6AS U.K.
3. Department of Botany, University of Sri Jayawardanapura; Nugegoda Sri Lanka
4. Institut für Botanik, Westfälische Wilhelms-Universität; D-48149 Münster Germany
5. Royal Botanic Garden Edinburgh; 20a Inverleith Row Edinburgh EH3 U.K