1. Ecology and Evolutionary Biology; University of Connecticut; 75 N Eagleville Road, Storrs, Connecticut 06269 U.S.A.
2. School of Botany and Zoology; University of Natal; Pietermaritzburg 3209 South Africa
3. Instituto Multidisciplinario de Biología Vegetal; CONICET-UNC, C. C 495 Córdoba 5000 Argentina