1. Graduate School of Biomedical Sciences, Hiroshima University, 1-2-3 Kasumi, Minami-ku, Hiroshima 734-8551, Japan
2. Ophthalmic Research & Development Center, Santen Pharmaceutical Co., Ltd, 8916–16 Takayama-cho, Ikoma, Nara 630-0101, Japan
3. Quantitative Biology Center, RIKEN, 6-2-3 Furuedai, Suita, Osaka 565-0871, Japan
4. Graduate School of Biomedical Sciences, Hiroshima University, 1-2-3 Kasumi, Minami-ku, Hiroshima 734-8551, Japan.