1. Institute of Translational Medicine, University of Liverpool, Ashton Street, Liverpool, L69 3GE, UK
2. Department of Colorectal Surgery, Countess of Chester Hospital NHS Foundation Trust, Chester, Cheshire, CH2 1UL, UK
3. Liverpool Hepatobiliary Unit, Aintree University Hospital, Longmoor Lane, Liverpool, Merseyside, L9 7AL, UK
4. Clatterbridge Cancer Centre, Bebington, Wirral, Merseyside, CH63 4JY, UK