1. Plant Protection Research Institute, Guangdong Academy of Agricultural Sciences, Guangzhou, Guangdong 510640, China; E-mails: joanyao0603@126.com (Q. Y.); linfa_quan2016@126.com (L. Q.); modeyouxiang@163.com (H. X.); 304087281@qq.com (T. J.); lwj4631