1. Key Laboratory of Adaptation and Evolution of Plateau Biota (AEPB), Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining 810008, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. Qinghai Province Key Laboratory of Crop Molecular Breeding, Xining 810008, China
4. Triticeae Research Institute, Sichuan Agricultural University, Chengdu 611130, China