1. School of Geography & Geosciences, University of St Andrews, North Street, St Andrews KY16 9LA, UK
2. Scottish Association of Marine Sciences, Scottish Marine Institute, Oban, PA37 1QA, UK
3. Department of Geography, College of Science, Swansea University, Singleton Park, Swansea SA2 8PP, UK
4. Godwin Laboratory for Paleoclimate Research, Department of Earth Sciences, University of Cambridge, Downing Street, Cambridge CB2 3EQ, UK