1. School of Human and Environmental Science, The University of Reading, Whiteknights, PO Box 217, Reading, RG6 6AH, UK
2. School of Ocean and Earth Science, National Oceanography Centre, University of Southampton, European Way, Southampton, SO14 3ZH, UK
3. Evolutionary Organismal Biology, Department of Physiology and Developmental Biology, Norbyvägen 18A, SE-752 36 Uppsala, Sweden
4. School of Earth and Ocean Sciences, Cardiff University, Main Building, Park Place, Cardiff CF10 3YE, Wales