1. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China
2. School of Earth Sciences, University of Bristol, Bristol BS8 1RJ, UK
3. School of GeoSciences, Yangtze University, Wuhan 430100, China