1. Department of Marine Sciences, University of the AegeanUniversity Hill, Mytilene 81100, Greece
2. School of Ocean and Earth Science, University of Southampton, Southampton Oceanography Centre, European WaySouthampton SO14 3ZH, UK (email: mbc@noc.soton.ac.uk)
3. Marine Research Division, AZTI Tecnalia, Herrera KaiaPortu aldea z/g, Pasaia 20110, Gipuzkoa, Spain
4. HR WallingfordHowbery Park, Wallingford OX10 8BA, UK