1. Molecular Neuroscience Research Center, Shiga University of Medical Science, Otsu, Japan
2. Department of Otolaryngology-Head and Neck Surgery, Kyoto Prefectural University of Medicine, Kyoto, Japan
3. Department of Physiology, School of Medicine, Kyusyu University, Fukuoka, Japan
4. Signaling Molecules Research Laboratory, National Institute of Advanced Industrial Science and Technology, Ibaraki, Japan
5. and Laboratory of Molecular Biology, Gifu Pharmaceutical University, Gifu, Japan