1. School of Marine Biosciences, Kitasato University, Sagamihara, Kanagawa 252-0373, Japan.
2. Faculty of Marine Bioscience, Fukui Prefectural University, Obama, Fukui 917-0003, Japan.
3. Fisheries Laboratory, The University of Tokyo, Hamamatsu, Shizuoka 431-0214, Japan.