1. University of Groningen, GELIFES (Groningen Institute for Evolutionary Life Sciences), Nijenborgh 7, 9747 AG Groningen, The Netherlands.
2. Institute of Biology, Molecular Evolution and Systematics of Animals, University of Leipzig, Talstraße 33, 04103 Leipzig, Germany.
3. Reptile, Amphibian and Fish Conservation Netherlands (RAVON), PO Box 1413, 5601 BK Nijmegen, The Netherlands.