1. Division of Environmental Dynamics, Graduate School of Environmental Science, The University of Shiga Prefecture, 2500 Hassaka, Hikone, Shiga 522-8533, Japan.
2. Institute of Systematics and Ecology of Animals, Siberian Branch of the Russian Academy of Sciences, 11 Frunze Street, Novosibirsk 630091, Russia.
3. Department of Ecosystem Studies, Faculty of Environmental Science, The University of Shiga Prefecture, 2500 Hassaka, Hikone, Shiga 522-8533, Japan.