1. Auburn University, Aquatic Parasitology Laboratory and Southeastern Cooperative Fish Parasite and Disease Laboratory, School of Fisheries, Aquaculture, & Aquatic Sciences, College of Agriculture, Auburn University, 203 Swingle Hall, Auburn, Alabama 3
2. African Amphibian Conservation Research Group, Unit for Environmental Sciences and Management, North-West University, Private Bag X6001, Potchefstroom, 2520, South Africa.
3. South African Institute for Aquatic Biodiversity, Somerset Street, Grahamstown 6139, South Africa.
4. Warm Springs Fish Technology Center, U.S. Fish and Wildlife Service, Auburn, Alabama 36849.