1. Centre for Sustainable Tropical Fisheries and Aquaculture, School of Marine and Tropical Biology, James Cook University, Cairns, Queensland 4878, Australia.
2. Fish Health Laboratory, Department of Fisheries Western Australia, South Perth, Western Australia 6151, Australia.
3. Centre for Biodiscovery and Molecular Development of Therapeutics, School of Public Health and Tropical Medicine, Queensland Tropical Health Alliance, James Cook University, Cairns, Queensland 4878, Australia.
4. Department of Veterinary Medicine, Cambridge Veterinary School, University of Cambridge, Cambridge CB3 0ES, U.K.
5. Parasitology and Marine Biodiversity, Biodiversity and Geosciences Program, Queensland Museum, South Brisbane, Queensland 4101, Australia.