1. Department of Geology and Paleontology, National Museum of Nature and Science, 4-1-1 Amakubo, Tsukuba, Ibaraki 305-0005, Japan (e-mail: )
2. Toko 15-4-2-3, Asahikawa, Hokkaido 078-8355, Japan
3. Higashi 3-11-2-25, Asahikawa, Hokkaido 070-0023, Japan